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In this blog we
will begin the discourse on ecology at the ecosystem level for reasons already
indicated;it is the level of greatest interest to everybody, regardless of
whether or not the subject of ecology is to be pursued beyond beyond the
introductory level. How, then, do we begin with something so formidable as an
ecological system. We being just as we would begin the study of any level – by
describing simplified versions, which encompass only the most important or
basic properties and functions. Since, in science, simplified versions of the
real world are called models, it would be appropriate at this point to talk a
little about models.
A model is a simplified formulation that mimics real – world
phenomena so that complex situations can be comprehended and predictions made.
In simplest form, models may be verbal or graphic, that is, consist of concise
statements or picture graphs. Although for the must part we shall restrict
ourselves to such ‘’informal’’ models in this book, it is important that we
consider at least the rationale of constructing more ‘’formal’’ models, because
such formulations are going to play an increasing role in decision – making
regarding man’s impact on his natural environment.
In its formal version a working model of an ecological
situation would, in most cases, have
four components, as listed below (with certain technical terms, as used by
systems analysts in parentheses).
1.
Properties (state variables)
2.
Forces (forcing functions), which are outside
energy sources or casual forces that drive the system.
3.
Flow pathways where energy flows or materials
transfers connect properties with each other and with forces.
4.
Interactions (interaction functions ) where
forces and properties interact to modify, amplify, or control flows.
Picture
A ayestem diagram showing four basic components of primary
interest in modelling ecosystems. See text for explanation.
Figure 1-2 shows how these components can be linked together
in a model diagram designed to mimic real – world situation. Shown are two
properties p1 and p2 which interact at I to produce or affect a third property
p3 when the system is driven by forcing E. five folow pathways are shown, with
F1 representing the input and F5 the output for the system as a whole.
The diagram could serve as a model for photochemical smog
production in the air over los angeles. In this case P1 could repre sent
hydrocarbons and P2 nitrogen oxides, two products of automobile exhaust
emission. Under the driving force of sunlight energy E, these interact to produce photochemical among P3 is
a more serious pollutant for man than is P1 and P2 acting alone.
Alternatively, figure 1-2 could represent a grassland
ecosystem in which P1 are the green plants, which convert sun energy E to food.
P2 might represent an herbivorous animal that eats plants, and P3 an omniverous
animal that can eat either the herbiveres or the plants. In this case the
interaction function I could represent several possibilities. It could be a
‘’no preference’’ switch if observations in the real world showed that the
omnivore P3 eats either P1 or P2 according to availability. Or I could be
specified to be a constant percentage value if it was found that the diet of P3
was composed of , say 80 percent plant and 20 persent animal matter
irrespective of the state of P1 and P2. Or I could be a seasonal switch if P3
feeds on plants during one part of the year and animals during another season.
Or it could e a threshold switch if P3 greatly prefers animal food and switches
to plants only when P2 is reduced to a low level.